Birth - Wikipedia
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Birth is the act or process of bearing or bringing forth offspring, also referred to in technical contexts as parturition. In mammals, the process is ... Birth FromWikipedia,thefreeencyclopedia Jumptonavigation Jumptosearch Processofbearingoffspring Thisarticleisaboutbirthinmammalsandotheranimals.Forbirthinhumans,seeChildbirth.Forotheruses,seeBirth(disambiguation). "Parturition"redirectshere.FortheStarTrekepisode,seeParturition(StarTrek:Voyager). Lambing:themotherlicksthefirstlambwhilegivingbirthtothesecond Birthistheactorprocessofbearingorbringingforthoffspring,[1]alsoreferredtointechnicalcontextsasparturition.Inmammals,theprocessisinitiatedbyhormoneswhichcausethemuscularwallsoftheuterustocontract,expellingthefetusatadevelopmentalstagewhenitisreadytofeedandbreathe. Insomespeciestheoffspringisprecocialandcanmovearoundalmostimmediatelyafterbirthbutinothersitisaltricialandcompletelydependentonparenting. Inmarsupials,thefetusisbornataveryimmaturestageafterashortgestationalperiodanddevelopsfurtherinitsmother'swomb'spouch. Itisnotonlymammalsthatgivebirth.Somereptiles,amphibians,fishandinvertebratescarrytheirdevelopingyounginsidethem.Someoftheseareovoviviparous,withtheeggsbeinghatchedinsidethemother'sbody,andothersareviviparous,withtheembryodevelopinginsideherbody,asinthecaseofmammals. Contents 1Mammals 1.1Humanchildbirth 1.2Cattle 1.3Dogs 1.4Marsupials 2Otheranimals 3Seealso 4References 5Citedtexts Mammals[edit] Largemammals,suchasprimates,cattle,horses,someantelopes,giraffes,hippopotamuses,rhinoceroses,elephants,seals,whales,dolphins,andporpoises,generallyarepregnantwithoneoffspringatatime,althoughtheymayhavetwinormultiplebirthsonoccasion. Intheselargeanimals,thebirthprocessissimilartothatofahuman,thoughinmosttheoffspringisprecocial.Thismeansthatitisborninamoreadvancedstatethanahumanbabyandisabletostand,walkandrun(orswiminthecaseofanaquaticmammal)shortlyafterbirth.[2] Inthecaseofwhales,dolphinsandporpoises,thesinglecalfisnormallyborntailfirstwhichminimizestheriskofdrowning.[3]Themotherencouragesthenewborncalftorisetothesurfaceofthewatertobreathe.[4] Mostsmallermammalshavemultiplebirths,producinglittersofyoungwhichmaynumbertwelveormore.Intheseanimals,eachfetusissurroundedbyitsownamnioticsacandhasaseparateplacenta.Thisseparatesfromthewalloftheuterusduringlaborandthefetusworksitswaytowardsthebirthcanal.[citationneeded] Largemammalswhichgivebirthtotwinsismuchmorerare,butitdoesoccuroccasionallyevenformammalsaslargeaselephants.InApril2018,approximately8-montholdelephanttwinsweresightedjoiningtheirmother'sherdintheTarangireNationalParkofTanzania,estimatedtohavebeenborninAugust2017.[5] Humanchildbirth[edit] Mainarticle:Childbirth Furtherinformation:Adaptationtoextrauterinelife Anillustrationofnormalhead-firstpresentationbytheobstetricianWilliamSmelliefromabout1792.Themembraneshaverupturedandthecervixisfullydilated. Humansusuallyproduceasingleoffspringatatime.Themother'sbodyispreparedforbirthbyhormonesproducedbythepituitarygland,theovaryandtheplacenta.[2]Thetotalgestationperiodfromfertilizationtobirthisnormallyabout38weeks(birthusuallyoccurring40weeksafterthelastmenstrualperiod).Thenormalprocessofchildbirthtakesseveralhoursandhasthreestages.Thefirststagestartswithaseriesofinvoluntarycontractionsofthemuscularwallsoftheuterusandgradualdilationofthecervix.Theactivephaseofthefirststagestartswhenthecervixisdilatedmorethanabout4 cmindiameterandiswhenthecontractionsbecomestrongerandregular.Thehead(orthebuttocksinabreechbirth)ofthebabyispushedagainstthecervix,whichgraduallydilatesuntilisfullydilatedat10 cmdiameter.Atsometime,theamnioticsacburstsandtheamnioticfluidescapes(alsoknownasruptureofmembranesorbreakingthewater).[6]Instagetwo,startingwhenthecervixisfullydilated,strongcontractionsoftheuterusandactivepushingbythemotherexpelsthebabyoutthroughthevagina,whichduringthisstageoflabouriscalledabirthcanalasthispassagecontainsababy,andthebabyisbornwithumbilicalcordattached.[7]Instagethree,whichbeginsafterthebirthofthebaby,furthercontractionsexpeltheplacenta,amnioticsac,andtheremainingportionoftheumbilicalcordusuallywithinafewminutes.[8] Enormouschangestakeplaceinthenewborn'scirculationtoenablebreathinginair.Intheuterus,theunbornbabyisdependentoncirculationofbloodthroughtheplacentaforsustenanceincludinggaseousexchangeandtheunbornbaby'sbloodbypassesthelungsbyflowingthroughtheforamenovale,whichisaholeintheseptumdividingtherightatriumandleftatrium.Afterbirththeumbilicalcordisclampedandcut,thebabystartstobreatheair,andbloodfromtherightventriclestartstoflowtothelungsforgaseousexchangeandoxygenatedbloodreturnstotheleftatrium,whichispumpedintotheleftventricle,andthenpumpedintothemainarterialsystem.Asaresultofthesechanges,thebloodpressureintheleftatriumexceedsthepressureintherightatrium,andthispressuredifferenceforcestheforamenovaletocloseseparatingtheleftandrightsidesoftheheart.Theumbilicalvein,umbilicalarteries,ductusvenosusandductusarteriosusarenotneededforlifeinairandintimethesevesselsbecomeligaments(embryonicremnants).[9] Cattle[edit] Acowgivingbirth Birthingincattleistypicalofalargermammal.Acowgoesthroughthreestagesoflaborduringnormaldeliveryofacalf.Duringstageone,theanimalseeksaquietplaceawayfromtherestoftheherd.Hormonechangescausesofttissuesofthebirthcanaltorelaxasthemother'sbodypreparesforbirth.Thecontractionsoftheuterusarenotobviousexternally,butthecowmayberestless.Shemayappearagitated,alternatingbetweenstandingandlyingdown,withhertailslightlyraisedandherbackarched.Thefetusispushedtowardthebirthcanalbyeachcontractionandthecow'scervixgraduallybeginstodilate.Stageonemaylastseveralhours,andendswhenthecervixisfullydilated.Stagetwocanbeseentobeunderwaywhenthereisexternalprotrusionoftheamnioticsacthroughthevulva,closelyfollowedbytheappearanceofthecalf'sfronthoovesandheadinafrontpresentation(oroccasionallythecalf'stailandrearendinaposteriorpresentation).[10]Duringthesecondstage,thecowwillusuallyliedownonhersidetopushandthecalfprogressesthroughthebirthcanal.Thecompletedeliveryofthecalf(orcalvesinamultiplebirth)signifiestheendofstagetwo.Thecowscramblestoherfeet(iflyingdownatthisstage),turnsroundandstartsvigorouslylickingthecalf.Thecalftakesitsfirstfewbreathsandwithinminutesisstrugglingtorisetoitsfeet.Thethirdandfinalstageoflaboristhedeliveryoftheplacenta,whichisusuallyexpelledwithinafewhoursandisofteneatenbythenormallyherbivorouscow.[10][11] Dogs[edit] Furtherinformation:Caninereproduction§ Gestationandlitters Birthistermedwhelpingindogs.[12]Amongdogs,aswhelpingapproaches,contractionsbecomemorefrequent.Labourinthebitchcanbedividedinto3stages.Thefirststageiswhenthecervixdilates,thiscausesdiscomfortandrestlessnessinthebitch.Commonsignsofthisstagearepanting,fasting,and/orvomiting.Thismaylastupto12hrs.[12]Stagetwoisthepassageoftheoffspring.[12]Theamnioticsaclookinglikeaglisteninggreyballoon,withapuppyinside,ispropelledthroughthevulva.Afterfurthercontractions,thesacisexpelledandthebitchbreaksthemembranesreleasingclearfluidandexposingthepuppy.Themotherchewsattheumbilicalcordandlicksthepuppyvigorously,whichstimulatesittobreathe.Ifthepuppyhasnottakenitsfirstbreathwithinaboutsixminutes,itislikelytodie.Furtherpuppiesfollowinasimilarwayonebyoneusuallywithlessstrainingthanthefirstusuallyat15-60minintervals.Ifapuphasnotbeenpassedin2hrsaveterinarianshouldbecontacted.[12]Stagethreeisthepassingoftheplacentas.Thisoftenoccursinconjunctionwithstagetwowiththepassingofeachoffspring.[12]Themotherwillthenusuallyeattheafterbirth.[13]Thisisanadaptiontokeepthedencleanandpreventitsdetectionbypredators.[12] Marsupials[edit] Seealso:Marsupial§ Reproductivesystem,andMarsupial§ Earlydevelopment Akangaroojoeyfirmlyattachedtoanippleinsidethepouch Aninfantmarsupialisborninaveryimmaturestate.[14]Thegestationperiodisusuallyshorterthantheintervalsbetweenoestrusperiods.Thefirstsignthatabirthisimminentisthemothercleaningoutherpouch.Whenitisborn,theinfantispink,blind,furlessandafewcentimetreslong.Ithasnostrilsinordertobreatheandforelegstoclingontoitsmother'shairsbutitshindlegsareundeveloped.Itcrawlsthroughitsmother'sfurandmakesitswayintothepouch.Hereitfixesontoateatwhichswellsinsideitsmouth.Itstaysattachedtotheteatforseveralmonthsuntilitissufficientlydevelopedtoemerge.[15]Joeysarebornwith"oralshields";inspecieswithoutpouchesorwithrudimentarypouchesthesearemoredevelopedthaninformswithwell-developedpouches,implyingaroleinmaintainingtheyoungattachedtothemother'snipple.[16] Otheranimals[edit] ACladoceragivingbirth(100xmagnification) Manyreptilesandthevastmajorityofinvertebrates,mostfish,amphibiansandallbirdsareoviparous,thatis,theylayeggswithlittleornoembryonicdevelopmenttakingplacewithinthemother.Inaquaticorganisms,fertilizationisnearlyalwaysexternalwithspermandeggsbeingliberatedintothewater(anexceptionissharksandrays,whichhaveinternalfertilization[17]).Millionsofeggsmaybeproducedwithnofurtherparentalinvolvement,intheexpectationthatasmallnumbermaysurvivetobecomematureindividuals.Terrestrialinvertebratesmayalsoproducelargenumbersofeggs,afewofwhichmayavoidpredationandcarryonthespecies.Somefish,reptiles,andamphibianshaveadoptedadifferentstrategyandinvesttheireffortinproducingasmallnumberofyoungatamoreadvancedstagewhicharemorelikelytosurvivetoadulthood.Birdscarefortheiryounginthenestandprovidefortheirneedsafterhatchinganditisperhapsunsurprisingthatinternaldevelopmentdoesnotoccurinbirds,giventheirneedtofly.[18] Ovoviviparityisamodeofreproductioninwhichembryosdevelopinsideeggsthatremaininthemother'sbodyuntiltheyarereadytohatch.Ovoviviparousanimalsaresimilartoviviparousspeciesinthatthereisinternalfertilizationandtheyoungareborninanadvancedstate,butdifferinthatthereisnoplacentalconnectionandtheunbornyoungarenourishedbyeggyolk.Themother'sbodyprovidesgasexchange(respiration),butthatislargelynecessaryforoviparousanimalsaswell.[18]Inmanysharkstheeggshatchintheoviductwithinthemother'sbodyandtheembryosarenourishedbytheegg'syolkandfluidssecretedbyglandsinthewallsoftheoviduct.[19]TheLamniformesharkspracticeoophagy,wherethefirstembryostohatchconsumetheremainingeggsandsandtigersharkpupscannibalisticallyconsumeneighbouringembryos.Therequiemsharksmaintainaplacentallinktothedevelopingyoung,thispracticeisknownasviviparity.Thisismoreanalogoustomammaliangestationthantothatofotherfishes.Inallthesecases,theyoungarebornaliveandfullyfunctional.[20]Themajorityofcaeciliansareovoviviparousandgivebirthtoalreadydevelopedoffspring.Whentheyounghavefinishedtheiryolksacstheyfeedonnutrientssecretedbycellsliningtheoviductandeventhecellsthemselveswhichtheyeatwithspecialistscrapingteeth.[21]TheAlpinesalamander(Salamandraatra)andseveralspeciesofTanzaniantoadinthegenusNectophrynoidesareovoviviparous,developingthroughthelarvalstageinsidethemother'soviductandeventuallyemergingasfullyformedjuveniles.[22] Amoredevelopedformofviviparitycalledplacentalviviparityisadoptedbysomespeciesofscorpions[23]andcockroaches,[24]certaingeneraofsharks,snakesandvelvetworms.Inthese,thedevelopingembryoisnourishedbysomeformofplacentalstructure.Theearliestknownplacentawasfoundrecentlyinagroupofextinctfishescalledplacoderms.AfossilfromAustralia'sGogoFormation,laiddownintheDevonianperiod,380 millionyearsago,wasfoundwithanembryoinsideitconnectedbyanumbilicalcordtoayolksac.Thefindconfirmedthehypothesisthatasub-groupofplacoderms,calledptyctodontids,fertilizedtheireggsinternally.Somefishesthatfertilizetheireggsinternallyalsogivebirthtoliveyoung,asseenhere.Thisdiscoverymovedourknowledgeoflivebirthback200 millionyears.[25]Thefossilofanothergenuswasfoundwiththreeembryosinthesameposition.[26]Placodermsareasistergroupoftheancestorofalllivingjawedfishes(Gnathostomata),includingbothchondrichthyians,thesharks&rays,andOsteichthyes,thebonyfishes. Amonglizards,theviviparouslizardZootocavivipara,slowwormsandmanyspeciesofskinkareviviparous,givingbirthtoliveyoung.SomeareovoviviparousbutotherssuchasmembersofthegeneraTiliquaandCorucia,givebirthtoliveyoungthatdevelopinternally,derivingtheirnourishmentfromamammal-likeplacentaattachedtotheinsideofthemother'suterus.Inarecentlydescribedexample,anAfricanspecies,Trachylepisivensi,hasdevelopedapurelyreptilianplacentadirectlycomparableinstructureandfunctiontoamammalianplacenta.[27]Viviparyisrareinsnakes,butboasandvipersareviviparous,givingbirthtoliveyoung. Femaleaphidgivingbirth Themajorityofinsectslayeggsbutaveryfewgivebirthtooffspringthatareminiatureversionsoftheadult.[18]Theaphidhasacomplexlifecycleandduringthesummermonthsisabletomultiplywithgreatrapidity.Itsreproductionistypicallyparthenogeneticandviviparousandfemalesproduceunfertilizedeggswhichtheyretainwithintheirbodies.[28]Theembryosdevelopwithintheirmothers'ovariolesandtheoffspringareclonesoftheirmothers.Femalenymphsarebornwhichgrowrapidlyandsoonproducemorefemaleoffspringthemselves.[29]Insomeinstances,thenewbornnymphsalreadyhavedevelopingembryosinsidethem.[18] Seealso[edit] Wikiquotehasquotationsrelatedto:Birth Animalsexualbehaviour Breedingseason Caesareansection Dystocia Episiotomy Foaling(horses) Forcepsdelivery Kegelexercises Matingsystem Odondevice Perinealmassage Reproduction Reproductivesystem Ventouse Birthspacing References[edit] ^"birth".OEDOnline.June2013.OxfordUniversityPress.Entry19395(accessed30August2013). ^abDorit,R.L.;Walker,W.F.;Barnes,R.D.(1991).Zoology.SaundersCollegePublishing.pp. 526–527.ISBN 978-0-03-030504-7. ^MarkSimmonds,WhalesandDolphinsoftheWorld,NewHollandPublishers(2007),Ch.1,p.32ISBN 1845378202. ^Crockett,Gary(2011)."HumpbackWhaleCalves".HumpbackwhalesAustralia.Archivedfromtheoriginalon2017-02-27.Retrieved2013-08-28. ^"TrunkTwins :ElephantTwinsBorninTarangire|AsiliaAfrica".AsiliaAfrica.2018-04-06.Retrieved2018-04-06. ^NICE(2007).Section1.6,Normallabour:firststage ^NICE(2007).Section1.7,Normallabour:secondstage ^NICE(2007).Section1.8,Normallabour:thirdstage ^Houston,Rob(editor);Lea,Maxine(arteditor)(2007).TheHumanBodyBook.DorlingKindersley.p. 215.ISBN 978-1-8561-3007-3.{{citebook}}:|author1=hasgenericname(help) ^ab"Calving".Alberta:AgricultureandRuralDevelopment.2000-02-01.Retrieved2013-08-28. ^"CalvingManagementinDairyHerds:TimingofInterventionandStillbirth"(PDF).TheOhioStateUniversityCollegeofVeterinaryMedicineExtension.2012.Retrieved2013-12-17. ^abcdefKustritz,M.(2005)."Reproductivebehaviourofsmallanimals".Theriogenology.64(3):734–746.doi:10.1016/j.theriogenology.2005.05.022.PMID 15946732. ^Dunn,T.J."Whelping:NewPuppiesOnTheWay!".PuppyCenter.PetMD.Retrieved2013-08-28. ^HughTyndale-Biscoe;MarilynRenfree(30January1987).ReproductivePhysiologyofMarsupials.CambridgeUniversityPress.ISBN 978-0-521-33792-2. ^"Reproductionanddevelopment".ThylacineMuseum.Retrieved2013-08-28. ^YvetteSchneiderNanette(Aug2011)."Thedevelopmentoftheolfactoryorgansinnewlyhatchedmonotremesandneonatemarsupials".J.Anat.219(2):229–242.doi:10.1111/j.1469-7580.2011.01393.x.PMC 3162242.PMID 21592102. ^SeaWorld,Sharks&RaysArchived2013-11-10attheWaybackMachine;accessed2013.09.09. ^abcdAttenborough,David(1990).TheTrialsofLife.pp. 26–30.ISBN 9780002199124. ^Adams,KyeR.;Fetterplace,LachlanC.;Davis,AndrewR.;Taylor,MatthewD.;Knott,NathanA.(January2018)."Sharks,raysandabortion:Theprevalenceofcapture-inducedparturitioninelasmobranchs".BiologicalConservation.217:11–27.doi:10.1016/j.biocon.2017.10.010.Archivedfromtheoriginalon2019-02-23.Retrieved2019-06-30. ^"Birthandcareofyoung".Animals:Sharksandrays.BuschEntertainmentCorporation.ArchivedfromtheoriginalonAugust3,2013.Retrieved2013-08-28. ^Stebbins,RobertC.;Cohen,NathanW.(1995).ANaturalHistoryofAmphibians.PrincetonUniversityPress.pp. 172–173.ISBN 978-0-691-03281-8. ^Stebbins,RobertC.;Cohen,NathanW.(1995).ANaturalHistoryofAmphibians.PrincetonUniversityPress.p. 204.ISBN 978-0-691-03281-8. ^Capinera,JohnL.,Encyclopediaofentomology.SpringerReference,2008,p.3311. ^Costa,JamesT.,TheOtherInsectSocieties.BelknapPress,2006,p.151. ^Dennis,Carina(2008-05-28)."NatureNews:Theoldestpregnantmum:Devonianfossilizedfishcontainsanembryo".Nature.453(7195):575.Bibcode:2008Natur.453..575D.doi:10.1038/453575a.PMID 18509405. ^Long,JohnA.;Trinastic,Kate;Young,GavinC.;Senden,Tim(2008-05-28)."LivebirthintheDevonianperiod".Nature.453(7195):650–652.Bibcode:2008Natur.453..650L.doi:10.1038/nature06966.PMID 18509443.S2CID 205213348. ^BlackburnDG,FlemmingAF(2012)."InvasiveimplantationandintimateplacentalassociationsinaplacentotrophicAfricanlizard,Trachylepisivensi(scincidae)".J.Morphol.273(2):137–59.doi:10.1002/jmor.11011.PMID 21956253.S2CID 5191828. ^Blackman,RogerL(1979)."Stabilityandvariationinaphidclonallineages".BiologicalJournaloftheLinneanSociety.11(3):259–277.doi:10.1111/j.1095-8312.1979.tb00038.x.ISSN 1095-8312. ^Conrad,Jim(2011-12-10)."Theaphidlifecycle".TheBackyardNatureWebsite.Retrieved2013-08-31. Citedtexts[edit] "Intrapartumcare:Careofhealthywomenandtheirbabiesduringchildbirth".NICE.September2007.Archivedfromtheoriginalon2014-04-26. 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