The Calvin Cycle - Hyperphysics

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Carbon dioxide is captured in a cycle of reactions known as the Calvin cycle or the Calvin-Benson cycle after its discoverers. It is also known as just the ... TheCalvinCycle Clickonanypartofthediagramformoredetail. Plantsuseenergyfromthesunintinyenergyfactoriescalledchloroplasts.Usingchlorophyllintheprocessofphotosynthesis,theyconvertthesun'senergyintostorableforminorderedsugarmoleculessuchasglucose.Inthisway,carbondioxidefromtheairandwaterfromthesoilinamoredisorderedstatearecombinedtoformthemoreorderedsugarmolecules. CarbondioxideiscapturedinacycleofreactionsknownastheCalvincycleortheCalvin-Bensoncycleafteritsdiscoverers.ItisalsoknownasjusttheC3cycle.ThoseplantsthatutilizejusttheCalvincycleforcarbonfixationareknownasC3plants.Carbondioxidediffusesintothestromaofchloroplastsandcombineswithafive-carbonsugar,ribulose1,5-biphosphate(RuBP).TheenzymethatcatalyzesthisreactionisreferredtoasRuBisCo,alargemoleculethatmaybethemostabundantorganicmoleculeontheEarth.Thiscatalyzedreactionproducesa6-carbonintermediatewhichdecaysalmostimmediatelytoformtwomoleculesofthe3-carboncompound3-phosphoglycericacid(3PGA).Thefactthatthis3-carbonmoleculeisthefirststableproductofphotosynthesisleadstothepracticeofcallingthiscycletheC3cycle. InC3plantsthephotosynthesis,carbonfixationandCalvincyclealloccurinasinglechloroplast. InC4plantsthephotosynthesistakesplaceinachloroplastofathin-walledmesophyllcellanda4-carbonacidishandedofftoathick-walledbundlesheathcellwheretheCalvincycleoccursinachloroplastofthatsecondcell.ThisprotectstheCalvincyclefromtheeffectsofphotorespiration. InCAMplantsthephotosynthesisandinitialcarbonfixationoccuratnightanda4-carbonacidisstoredinthecell'svacuole.Duringtheday,theCalvincycleoperatesinthesamechloroplasts. EnergyfromATPandfromthereducedcoenyzmeNADPHisusedtoremoveaphosphategroupfrom3PGAandreducetheresultingdiphosphoglycerate(DPGA)toproducethe3-carbonsugarglyceraladehyde-3-phosphate(G3P).SomeofthisG3PisusedtoregeneratetheRuBPtocontinuethecycle,butsomeisavailableformolecularsynthesisandisusedtomakefructosediphosphate.Thefructosediphosphateisthenusedtomakeglucose,sucrose,starchandothercarbohydrates. Energycycleinlivingthings IndexPhotosynthesisConceptsReferenceMoore,etal.Ch7  HyperPhysics*****BiologyRNave GoBack



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